Publications

Check out our publications on Google Scholar

Peer-reviewed

  1. Albery, G.F., Becker, D.J., Firth, J.A., De Moor, D., Ravindran, S., Silk, M., Sweeny, A.R., Vander Wal, E., Webber, Q., Allen, B., Babayan, S.A., Barve, S., Begon, M., Birtles, R.J., Block, T.A., Block, B.A., Bradley, J.E., Budischak, S., Buesching, C., Burthe, S.J., Carlisle, A.B., Caselle, J.E., Cattuto, C., Chaine, A.S., Chapple, T.K., Cheney, B.J., Clutton-Brock, T., Collier, M., Curnick, D.J., Delahay, R.J., Farine, D.R., Fenton, A., Ferretti, F., Feyrer, L., Fielding, H., **Foroughirad, V.**.. S. Bansal. (2025) Density-dependent network structuring within and across wild animal systems. *Nat Ecol Evol* 1–12. doi:10.1038/s41559-025-02843-z


  2. Collier, M.A., Jacoby, A., **Foroughirad, V.**, Patterson, E.M., Krzyszczyk, E., Wallen, M.M., Miketa, M.L., Karniski, C., Wilkin, S., Mann, J., & S. Bansal. (2025) Breathing synchrony shapes respiratory disease risk in bottlenose dolphins. *Communications Biology*, 8, 870. [doi:10.1038/s42003-025-08161-1](https://doi.org/10.1038/s42003-025-08161-1)


  3. Meier, A. C., Restrepo, O.N., Nordseth, A., Copeland, M., **Foroughirad, V.**, Mann, J., Wittemyer, G., J.E. Smith. (2025) Network indicators of cultural resilience to anthropogenic removals in animal societies. *Philosophical Transactions of the Royal Society B.* [doi:10.1098/rstb.2024.0144](https://doi.org/10.1098/rstb.2024.0144)


  4. Vivier, F., Andrés, C., Gonzalvo, J., Fertitta, K., van Aswegen, M., **Foroughirad, V.**, Mann, J., McEntee, M., Wells, R.S., & L. Bejder. (2024) Inferring dolphin population status: using unoccupied aerial systems to quantify age-structure. *Animal Conservation*. [doi:10.1111/acv.12978](https://doi.org/10.1111/acv.12978)


  5. McEntee, M.H.F., **Foroughirad, V.**, Krzyszczyk, E. & J. Mann. (2023) Sex bias in mortality risk varies with age and sex specific selection pressures across the lifespan of bottlenose dolphins. *Proceedings of the Royal Society B*. [doi:10.1098/rspb.2023.0675](https://doi.org/10.1098/rspb.2023.0675)


  6. Crystal-Ornelas, R., Edwards, B., Hébert, K., Hudgins, E.J., Sánchez-Reyes, L.L., Scott, E.R., Grainger, M., **Foroughirad, V.**, Binley, A., Brookson, C., Gaynor, K., Shafiei Sabet, S., Güncan, A., Hillemann, F., Weierbach, H., Gomes, D.G.E., Braga, P.H.P. (2023) Not just for programmers: How GitHub can accelerate collaborative and reproducible research in ecology and evolution. *Methods in Ecology and Evolution*. [doi:10.1111/2041-210X.14108](https://doi.org/10.1111/2041-210X.14108)


  7. Dolezal, M., **Foroughirad, V.**, Fish, F., Jacoby, A.M., Collier, M., Murphy, C., Rittmaster, K., Bansal, S., Mann, J. (2023) Some like it hot: Temperature and hydrodynamic factors influence Xenobalanus globicipitis attachment to cetaceans. *Marine Mammal Science*. [doi:10.1111/mms.13022](https://doi.org/doi:10.1111/mms.13022)


  8. **Foroughirad, V.**, Frère, C.H., Levengood, A.L., Kopps, A.M., Krzyszczyk, E. & J. Mann. (2023) Small effects of family size on sociality despite strong kin preferences in female bottlenose dolphins. *Animal Behaviour*. [doi:10.1016/j.anbehav.2022.10.011](https://doi.org/10.1016/j.anbehav.2022.10.011)


  9. Gomes, D.G.E., Pottier, P., Crystal-Ornelas, R., Hudgins, E.J., **Foroughirad, V.**, Sánchez-Reyes, L.L., Turba, R., Martinez, P.A., Moreau, D., Bertram, M., Smout, C. & K. Gaynor. (2022) Why don’t we share data and code? Perceived barriers and benefits to public archiving practices. *Proceedings of the Royal Society B.* [doi:10.1098/rspb.2022.1113](https://doi.org/10.1098/rspb.2022.1113)


  10. **Foroughirad, V.**, McEntee, M. H., Kopps, A.M., Levengood, A.L., Frère, C.H & J. Mann. (2022) Reproductive timing as an explanation for skewed parentage assignment ratio in a bisexually philopatric population. *Behavioral Ecology and Sociobiology*. 76, 129. [doi:10.1007/s00265-022-03233-2](https://doi.org/10.1007/s00265-022-03233-2)


  11. Levengood, A. L., Strickland, K., **Foroughirad, V.**, Mann, J., Cristescu, R.H., Krzyszczyk, E.B., Frère, C. (2022) Heterogeneity of resource competition shapes individual variation in dolphin social behavior. *Behavioral Ecology*. Volume 33, Issue 4, July/August 2022, Pages 745–757, [doi:10.1093/beheco/arac037](https://doi.org/10.1093/beheco/arac037)


  12. Strickland K., Mann J., **Foroughirad V.**, Levengood A.L., Frère C.H. (2021) Maternal effects and fitness consequences of individual variation in bottlenose dolphins’ ecological niche. *Journal of Animal Ecology*. 2021; 00:1–13. [doi:10.1111/1365-2656.13513](https://doi.org/10.1111/1365-2656.13513)


  13. Mann, J., **Foroughirad, V.**, McEntee, M.H., Miketa, M.L., Evans, T., Karniski, C.B., Krzyszczyk, E.B., Patterson, EM., Strohman, J., Wallen, M.M. (2021) Elevated calf mortality and long-term responses of wild bottlenose dolphins to extreme climate events: Impacts of foraging specialization and human provisioning. *Frontiers in Marine Science*. 8:617550. [doi:10.3389/fmars.2021.617550](https://doi.org/10.3389/fmars.2021.617550)


  14. Galezo, A., **Foroughirad, V.**, Krzyszczyk, E., Frère, C. & Mann, J. (2020). Juvenile social dynamics reflect adult reproductive strategies in bottlenose dolphins. *Behavioral Ecology*. [doi:10.1093/beheco/araa068](https://doi.org/10.1093/beheco/araa068)


  15. Powell, S.N., Wallen, M.M., Miketa, M.L., Krzyszczyk, E., **Foroughirad, V.**, Bansal, S. and Mann, J. (2019) Sociality and tattoo skin disease among bottlenose dolphins in Shark Bay, Australia. *Behavioral Ecology*. [doi:10.1093/beheco/arz207](https://doi.org/10.1093/beheco/arz207)


  16. **Foroughirad, V.**, Levengood, A., Mann, J. & Frère, C. (2019) Quality and quantity of genetic relatedness data affect the analysis of social structure. *Molecular Ecology Resources*. [doi:10.1111/1755-0998.13028](https://doi.org/10.1111/1755-0998.13028)


  17. Miketa, M., Patterson, EM., Krzyszczyk, E., **Foroughirad, V.**, & Mann, J. (2018) Calf age and sex affects maternal diving behavior in Shark Bay bottlenose dolphins. *Animal Behaviour*. [doi:10.1016/j.anbehav.2017.12.023](https://doi.org/10.1016/j.anbehav.2017.12.023)


  18. Strickland, K., Levengood, A., **Foroughirad, V.**, Mann, J., Krzyszczyk, E., & Frère, C. (2017) A framework for the identification of long-term social avoidance in longitudinal datasets. *Royal Society Open Science*. 4(8) [doi:10.1098/rsos.170641](https://doi.org/10.1098/rsos.170641)


  19. Karniski, C.B., Patterson, E.M., Krzyszczyk, E., **Foroughirad, V.**, Stanton, M.A. & Mann, J. (2015) A comparison of survey and focal follow methods for estimating individual activity budgets of cetaceans. *Marine Mammal Science*. [doi:10.1111/mms.12198](https://doi.org/10.1111/mms.12198)


  20. **Foroughirad, V.** & Mann, J. (2013) Long-term impacts of fish provisioning on the behavior and survival of bottlenose dolphins. *Biological Conservation*. 160:242-249. [doi:10.1016/j.biocon.2013.01.001](https://doi.org/10.1016/j.biocon.2013.01.001)

Preprints

Rankin, R.W., Foroughirad, V., Krzyszczyk, E.B., Frère, C.H., & J. Mann. Changes in social position predict survival in bottlenose dolphins. bioRxiv, 2022.08. 25.505273

Meehan, K.C., Class, B., Nakagawa, S., Foroughirad, V., Mann, J., Frere, C., 2025. The age of change: social aging in dolphins. EcoEvoRxiv, doi:10.32942/X2KW8W

Manlik, O., Mann, J., Krützen, M., Foroughirad, V., McEntee, M., Kopps, A., Frere, C., Allen, S., Connor, R., Bejder, L., Sherwin, W., 2025. MHC diversity confers transgenerational fitness advantage in wild dolphins. ResearchSquare, doi:10.21203/rs.3.rs-7279666